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dsff For Wales, centre Jonathan Davies has been recalled to the starting line-up after four months out of the side with a torn pectoral muscle.
Lock Alun Wyn Jones will also start having missed the victory over France due to a foot infection.
Fellow lock Luke Charteris was also due to start but has now been ruled out with a neck injury with Jake Ball, who made his debut in Wales' victory over France, taking his place in the starting line-up.
Scarlets' Liam Williams drops to the bench and the Dragons' Andrew Coombs comes into the replacements.
Gatland's line-up contains 11 of the team that ended England's Grand Slam hopes last season, with Ball, scrum-half Rhys Webb, fly-half Rhys Priestland and flanker Dan Lydiate the only players involved this time who did not start that game.
England: 15 Brown, 14 Nowell, 13 Burrell, 12 Twelvetrees, 11 May, 10 Farrell, 9 Care; 1 Marler, 2 Hartley, 3 Wilson, 4 Launchbury, 5 Lawes, 6 Wood, 7 Robshaw, 8 Morgan.
Replacements: 16 T Youngs, 17 M Vunipola, 18 Thomas, 19 Attwood, 20 Johnson, 21 Dickson, 22 Ford, 23 Goode.
Wales: 15 Halfpenny, 14 Cuthbert, 13 Davies, 12 Roberts, 11 North, 10 Priestland, 9 Webb; 1 Jenkins, 2 Hibbard, 3 A Jones, 4 Charteris, 5 Wyn Jones, 6 Lydiate, 7 Warburton, 8 Faletau.